Плио-плейстоценовые герпетофауны окрестностей села Коротояк (Воронежская область)
Ключевые слова:
плиоцен, плейстоцен, амфибии, рептилии, остеология, палеогеография, остеопатология
Аннотация
Введение: На юго-восточной окраине села Коротояк (Воронежская область) в трех местонахождениях разного возраста (Коротояк-1, Коротояк-3 и Коротояк-4) найдены остатки земноводных и пресмыкающихся. Целью статьи является информация о результатах определения этих остатков и их интерпретация. Систематическая часть. Приводятся описания и идентификация найденных костей амфибий и рептилий. Обсуждение и выводы. Всего две кости, идентифицированные как Pelophylax ridibundus и Natrix sp., происходят из местонахождения Коротояк-1 позднеплиоценового возраста. В раннеэоплейстоценовом местонахождении Коротояк-3 были определены Palaeobatrachus sp. – 1, Bufo sp. – 2, Anura indet. – 2, Lacertidae indet. – 1, Serpentes indet. – 2. Коллекция Коротояк-4 мучкапского возраста включает Bombina sp. – 1, Pelobates sp. – 5, Palaeobatrachus sp. – 1, Bufo verrucosissimus – 1, Bufo sp. – 3, Pelophylax ridibundus – 1, Rana temporaria – 2, Rana sp. – 1, Ranidae indet. – 3, Lacerta cf. agilis – 1, Elaphe dione – 2, Colubridae indet. – 1, Vipera sp. – 1. Остатки амфибий и рептилий во всех местонахождениях говорят о благоприятных условиях среды во время их формирования. Находки остатков Bufo sp. и Palaeobatrachus sp. свидетельствуют о сравнительно влажных лесных условиях во время формирования местонахождения Коротояк-3. Герпетофауна местонахождения Коротояк-4 позволяет предположить существование лиственного или смешанного леса и более теплый по сравнению с нынешним климат. В ее составе найдены современные виды, ареалы которых находятся вдалеке от местонахождения. Bufo verrucosissimus обитает ныне на Кавказе. Северная граница ареала Elaphe dione в настоящее время проходит через Ростовскую область. Сфенэтмоид палеобатрахуса найден на территории Восточно-Европейской равнины впервые. Этот экземпляр в деталях отличается от всех находок сфенэтмоидов в Западной и Центральной Европе. Однако из-за невозможности оценить изменчивость этого элемента скелета он определен только до родового уровня. Три фрагмента этой коллекции имеют патологические изменения: срастание позвонков у чесночницы и утолщение под котилярным ободом на позвонках узорчатого полоза и гадюки.
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Литература
1. Shik S. M., Tesakov A. S., Agadjanyan A. K., Iosifova Yu. I., Markova A. K., Pisareva V. V., Semenov V. V. Proekt regional'noi stratigraficheskoi shkaly eopleistocena i gelazia (paleopleistotsen) tsentra i yuga Evropeiskoi Rossii [Draft regional stratigraphic scale of the Eopleistocene and Gelazian (Paleopleistocene) of center and south of European Russia]. Bulleten' regional'noi mezhvedomstvennoi stratigraficheskoi komissii po Tsentru i yugu Russkoi platformy. [Bulletin of the Regional Interdepartmental Stratigraphic Commission for the Center and South of the Russian Platform], 2015, no. 6, pp. 97–107. (In Russ.)
2. Agadjanyan A. K. Melkie mlekopitayushchie pliotsen–pleistotsena Russkoi ravniny [Pliocene–Pleistocene Small Mammals of the Russian Plain]. Trudy Paleontologicheskogo Instituta RAN [The work of the Paleontological Institute of RAS], Moscow, Nauka, vol. 289, 2009, 676 p. (In Russ.)
3. Agadjanyan A. K., Глушанкова Н. И. Paleogeografiya pliotsen–pleistotsena verchnego Dona. [Paleogeography of the Pliocene Pleistocene of the Upper Don]. Bulletin of the Commission for Study of the Quaternary. Moscow, GIN, 1988, no. 57, pp. 62–77. (In Russ.)
4. Shik S. M., Agadjanyan A. K., Iosifova Yu. I., Markova A. K., Pisareva V. V., Semenov V. V., Tesakov A. S. Proekt regional'noi stratigraficheskoi shkaly eopleistocena tsentra i yuga Vostochno-Evropeiskoi platformy. [Draft regional stratigraphic scale of the Eopleistocene of the center and southeast of the East European platform]. Fundamental'nye problemy kvartera, itogi izucheniya i osnovnye napravleniya dal'neishikh issledovanii [Fundamental problems of quaternary, results and main trends of future studies], Irkutsk, September 15–20, 2015, p. 505–507. (In Russ.)
5. Ratnikov V. Yu. Beskhvostye amfibii pozdnego kainozoya Vostochno-Evropeiskoi platformy i ikh stratigraficheskoe i paleogeograficheskoe znachenie. [Tailless amphibians of the Late Cenozoic of the East European Platform and their stratigraphic and paleogeographic significance]. Voronezh, VINITI, 1994. 140 p. (In Russ.)
6. Szyndlar Z. Fossil snakes from Poland. Acta zool. cracov., 1984. vol. 28, no 1, pp. 1–156.
7. Evans S. E. The skull of lizards and Tuatara. Biology of the Reptilia, 2008, vol. 20, The skull of Lepidosauria, pp. 1–347.
8. Ratnikov V. Yu. K identifikatsii zherlianok (Bombina, Discoglossidae) po elementam skeleta pri paleontologicheskikh issledovaniyakh [To identification of toads (Bombina, Discoglossidae) by skeletal elements in paleontological studies] Aktual'nye problemy gerpetologii i toksinologii [Actual problems of herpetology and toxinology]. Tol'yatti, 2003, no. 6, pp. 91–101. (In Russ.)
9. Ratnikov V. Yu. Pozdnekainozoiskie zemnovodnye i cheshuichatye presmykayushchiesya Vostochno-Evropeiskoi ravniny. [Late Cenozoic amphibians and scaly reptiles of the East European Plain]. Trudy Nauchno-issledovatel'skogo Instituta Geologii [The work of the Research Institute of Geology], Voronezh, VSU Publ., vol. 10, 2002, 138 p. (In Russ.)
10. Villa A., Roček Z., Tschopp E., van den Hoek Ostende L. W., Delfino M. Palaeobatrachus eurydices, sp. nov. (Amphibia, Anura), the last Western European Palaeobatrachid. Journal of Vertebrate Paleontology. 2016, vol. 36, no. 6, pp. 1–15.
11. Špinar Z. Tertiary Frogs from Central Europe. Prague: Academia, 1972, 286 p.
12. Hossini S., Rage J.-C. Palaeobatrachid frogs from the earliest Miocene (Agenian) of France, with description of a new species. Geobios, 2000, vol. 33, no. 2, pp. 223–231.
13. Venczel M., Codrea V., Fărcaş C. A new palaeobatrachid frog from the early Oligocene of Suceag, Romania. Journal of Systematic Palaeontology, 2013, vol. 11, no 2, pp. 179–189.
14. Sanchiz B., Młynarski M. Remarks on the fossil anurans from the Polish Neogene. Acta zool. cracov., 1979, vol. 24, no. 1–4, pp. 153–173.
15. Wuttke M., Přikryl T., Ratnikov V. Yu., Dvořák Z., Roček Z. Generic diversity and distributional dynamics of the Palaeobatrachidae (Amphibia: Anura). Palaeobiodiversity and Palaeoenvironments, 2012, vol. 92, no. 3, pp. 367–395.
16. Ratnikov V. Yu. The first find of a Pliobatrachus (Anura, Palaeobatrachidae) in muchkapian sediments of the Don Basin. Paleont. Zhurn., 1993, no. 4, pp. 130–132. (In Russ.)
17. Ratnikov V. Yu. Rannepleistotsenovaya gerpetofsuna iz mestonakhozhdeniya Kozii Ovrag v basseine Verkhnego Dona. [Early Pleistocene herpetofauna from the Koziy Ovrag locality in the Upper Don basin]. Izv. vuzov. Geologiya i razvedka = Proceedings of universities. Geology and exploration, 1996, no. 6, pp. 16–19. (In Russ.)
18. Ratnikov V. Yu. The Late Pliocene herpetofauna of the Korotoyak site the Voronezh region and the contemporary lanscape. The down of the Quaternary: Abstracts of the IN-QUA-SEQS symposium, 16-21 June 1996, Kerkrade, The Netherlands, p. 79.
19. Ратников В. Ю. On the finds of Pliobatrachus (Anura, Palaeobatrachidae) in Eastern Europe. Paleintological Journal, 1997, vol. 31, no. 4, pp. 415–421.
20. Ratnikov V. Yu. Pliocene anurans of East-European platform. Russian journal of herpetology, 2001, vol.8, no 3, pp. 171–178.
21. Ratnikov V. Yu. Muchkapian (Early Neopleistocene) Amphibians and Reptiles of the East-European plain. Russian journal of herpetology, 2002, vol.9, no. 3, P. 229–236.
22. Ratnikov V. Yu. Novye nakhodki zemnovodnykh i и presmykayushchikhsya v opornykh muchkapskikh mestonakhozhdeniyakh basseina Verkhnego Dona [New finds of amphibians and reptiles in the reference Muchkap localities of the Upper Don basin]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya = Proceedings of Voronezh State University. Series: Geology, 2002, no. 1, pp. 73–79. (In Russ.)
23. Ratnikov V. Yu. Novye nakhodki zemnovodnykh i presmykayushchikhsya iz nizhneneopleistotsenovogo mestonakhozhdeniya Il'inka [New Finds of Amphibians and Reptiles from the Lower Neopleistocene Ilyinka Locality]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya = Proceedings of Voronezh State University. Series: Geology, 2003, no. 1, pp. 36–39. (In Russ.)
24. Ratnikov V. Yu. Osteology of Russian toads and frogs for palaeontological researches. Acta zool. cracov., 2001, vol. 44, no. 1, pp. 1–23.
25. Ratnikov V. Yu. Osteologicheskie kharakteristiki nadvidovykh taksonov zhab i lyagushek Vostochnoi Evropy [Osteological characteristics of supraspecific taxa of toads and frogs of Eastern Europe] Voprosy gerpetologii: materialy pyatogo s'ezda Gerpetologicheskogo obshchestva im. A.M. Nikol'ckogo [Questions of herpetology: materials of the Fifth Congress of the Herpetological Society named after A.M. Nikolsky], Minsk, September 25-28, 2012, pp. 269–273. (In Russ.)
26. Čerňansky ́ A., Syromyatnikova E. V. The first Miocene fossils of Lacerta cf. trilineata (Squamata, Lacertidae) with a comparative study of the main cranial osteological differences in green lizards and their relatives. PLoS ONE, 2019, vol.14, no. 8: e0216191, pp. 1–55.
27. Vorob'eva E. I., Chugunova T. Yu. Zubnaya sistema yashcherits: taksonomicheskoe i ekologicheskoe raznoobrazie. [Dental system of lizards: taxonomic and ecological diversity]. Moscow, Nauka, 1995, 151 p. (In Russ.)
28. Goncharov A. G. O rasprostranenii yashcherits (Sauria) v Tsentral'nom Chernozem'e. [About the distribution of lizards (Sauria) in the Central Black Earth Region]. Sovremennaya gerpetologiya: problemy i puti ikh resheniya. Stat'i po materialam dokladov Pervoi mezhdunarodnoi molodezhnoi konferentsii gerpetologov Rossii i sopredel'nykh stran. [Modern herpetology: problems and solutions. Articles based on the materials of the reports of the First International Youth Conference of Herpetologists in Russia and neighboring countries]. Saint-Peterburg, Russia, November 25–27, 2013, pp. 72–76. (In Russ.)
29. Ratnikov V. Yu. Gerpetofauna iz sredneneopleistotsenovogo mestonakhozhdeniya Ruchei Kolabinsky (Kursraya oblast'). [Herpetofauna from the Late Middle Pleistoсene locality of Kolabinsky brook (Kursk region). Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya = Proceedings of Voronezh State University. Series: Geology, 2019, no. 2, pp. 36–42. (In Russ.)
30. Szyndlar Z. A review of Neogene and Quaternary snakes of Central and Eastern Europe. Part II: Natricinae, Elapidae, Viperidae. Estudios geol., 1991, vol. 47, pp. 237–266.
31. Auffenberg W. The fossil Snakes of Florida. Tulane Studies Zool., 1963, vol. 10, no 3, pp. 131–216.
32. Ratnikov V. Yu. Identification of some Eurasian species of Elaphe (Colubridae, Serpentes) on the basis of vertebrae. Russian Journal of Herpetology, 2004, vol. 11, no. 2, pp. 91–98.
33. Ratnikov V., Mebert K. Fossil remains of Natrix tessellata from the Late Cenozoic deposits of the East European Plain. Mertensiella, 2011, no. 18, pp. 337–342.
34. Ananjeva N. B., Orlov N. L., Khalikov R. G., Darevsky I. S., Ryabov S. A., Barabanov A. V. The Reptiles of Northern Eurasia (Taxonomic Diversity, Distribution, Conservation Status). Saint-Petersburg, Zoological Institute RAS, 2006, 245 p.
35. Ivanov M. The first European pit viper from the Miocene of Ukraine. Acta Palaeontologica Polonica, 1999, vol. 44, no. 3, pp. 327–334.
36. Ratnikov V. Yu. Gerpetofauna nizhnepleistotsenovogo mestonakhozhdeniya Vol'naya Vershina v basseine Verkhnego Dona [Herpetofauna of the Lower Pleistocene locality Volnaya Vershina in the Upper Don basin]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya = Proceedings of Voronezh State University. Series: Geology, 1996, no. 1, pp. 90–93. (In Russ.)
37. Ratnikov V. Yu. Eopleistocene and Pleistocene Anurans of the East-European Platform. Paleont. Zhurn., 1992, no. 1, pp. 89– 100. (In Russ.)
38. Ratnikov V. Yu. Novye dannye o gerpetofaune mestonakhozhdeniya Kuznetsovka v Tambovskoi oblasti. [New data on the herpetofauna of the Kuznetsovka locality in the Tambov region]. Izv. vuzov. Geologiya i razvedka = Proceedings of universities. Geology and exploration, 1997, no. 1, pp. 26–32. (In Russ.)
39. Ratnikov V. Yu. Fossil remains of modern amphibian and reptile species as the material for studying of their areas history. Trudy Nauchno-issledovatel'skogo Instituta Geologii [The work of the Research Institute of Geology], Voronezh, VSU Publ., vol. 59, 2009, 91 p. (In Russ.).
40. Ratnikov V. Yu. Gerpetofauna verkhnepleistotsenovogo mestonakhozhdeniya Elasy v basseine Volgi. [Herpetofauna of the Upper Pleistocene Elasy locality in the Volga basin]. Aktual'nye problemy gerpetologii i toksinologii [Actual problems of herpetology and toxinology]. Tol'yatti, 2001, no. 5, pp. 81–88. (In Russ.)
2. Agadjanyan A. K. Melkie mlekopitayushchie pliotsen–pleistotsena Russkoi ravniny [Pliocene–Pleistocene Small Mammals of the Russian Plain]. Trudy Paleontologicheskogo Instituta RAN [The work of the Paleontological Institute of RAS], Moscow, Nauka, vol. 289, 2009, 676 p. (In Russ.)
3. Agadjanyan A. K., Глушанкова Н. И. Paleogeografiya pliotsen–pleistotsena verchnego Dona. [Paleogeography of the Pliocene Pleistocene of the Upper Don]. Bulletin of the Commission for Study of the Quaternary. Moscow, GIN, 1988, no. 57, pp. 62–77. (In Russ.)
4. Shik S. M., Agadjanyan A. K., Iosifova Yu. I., Markova A. K., Pisareva V. V., Semenov V. V., Tesakov A. S. Proekt regional'noi stratigraficheskoi shkaly eopleistocena tsentra i yuga Vostochno-Evropeiskoi platformy. [Draft regional stratigraphic scale of the Eopleistocene of the center and southeast of the East European platform]. Fundamental'nye problemy kvartera, itogi izucheniya i osnovnye napravleniya dal'neishikh issledovanii [Fundamental problems of quaternary, results and main trends of future studies], Irkutsk, September 15–20, 2015, p. 505–507. (In Russ.)
5. Ratnikov V. Yu. Beskhvostye amfibii pozdnego kainozoya Vostochno-Evropeiskoi platformy i ikh stratigraficheskoe i paleogeograficheskoe znachenie. [Tailless amphibians of the Late Cenozoic of the East European Platform and their stratigraphic and paleogeographic significance]. Voronezh, VINITI, 1994. 140 p. (In Russ.)
6. Szyndlar Z. Fossil snakes from Poland. Acta zool. cracov., 1984. vol. 28, no 1, pp. 1–156.
7. Evans S. E. The skull of lizards and Tuatara. Biology of the Reptilia, 2008, vol. 20, The skull of Lepidosauria, pp. 1–347.
8. Ratnikov V. Yu. K identifikatsii zherlianok (Bombina, Discoglossidae) po elementam skeleta pri paleontologicheskikh issledovaniyakh [To identification of toads (Bombina, Discoglossidae) by skeletal elements in paleontological studies] Aktual'nye problemy gerpetologii i toksinologii [Actual problems of herpetology and toxinology]. Tol'yatti, 2003, no. 6, pp. 91–101. (In Russ.)
9. Ratnikov V. Yu. Pozdnekainozoiskie zemnovodnye i cheshuichatye presmykayushchiesya Vostochno-Evropeiskoi ravniny. [Late Cenozoic amphibians and scaly reptiles of the East European Plain]. Trudy Nauchno-issledovatel'skogo Instituta Geologii [The work of the Research Institute of Geology], Voronezh, VSU Publ., vol. 10, 2002, 138 p. (In Russ.)
10. Villa A., Roček Z., Tschopp E., van den Hoek Ostende L. W., Delfino M. Palaeobatrachus eurydices, sp. nov. (Amphibia, Anura), the last Western European Palaeobatrachid. Journal of Vertebrate Paleontology. 2016, vol. 36, no. 6, pp. 1–15.
11. Špinar Z. Tertiary Frogs from Central Europe. Prague: Academia, 1972, 286 p.
12. Hossini S., Rage J.-C. Palaeobatrachid frogs from the earliest Miocene (Agenian) of France, with description of a new species. Geobios, 2000, vol. 33, no. 2, pp. 223–231.
13. Venczel M., Codrea V., Fărcaş C. A new palaeobatrachid frog from the early Oligocene of Suceag, Romania. Journal of Systematic Palaeontology, 2013, vol. 11, no 2, pp. 179–189.
14. Sanchiz B., Młynarski M. Remarks on the fossil anurans from the Polish Neogene. Acta zool. cracov., 1979, vol. 24, no. 1–4, pp. 153–173.
15. Wuttke M., Přikryl T., Ratnikov V. Yu., Dvořák Z., Roček Z. Generic diversity and distributional dynamics of the Palaeobatrachidae (Amphibia: Anura). Palaeobiodiversity and Palaeoenvironments, 2012, vol. 92, no. 3, pp. 367–395.
16. Ratnikov V. Yu. The first find of a Pliobatrachus (Anura, Palaeobatrachidae) in muchkapian sediments of the Don Basin. Paleont. Zhurn., 1993, no. 4, pp. 130–132. (In Russ.)
17. Ratnikov V. Yu. Rannepleistotsenovaya gerpetofsuna iz mestonakhozhdeniya Kozii Ovrag v basseine Verkhnego Dona. [Early Pleistocene herpetofauna from the Koziy Ovrag locality in the Upper Don basin]. Izv. vuzov. Geologiya i razvedka = Proceedings of universities. Geology and exploration, 1996, no. 6, pp. 16–19. (In Russ.)
18. Ratnikov V. Yu. The Late Pliocene herpetofauna of the Korotoyak site the Voronezh region and the contemporary lanscape. The down of the Quaternary: Abstracts of the IN-QUA-SEQS symposium, 16-21 June 1996, Kerkrade, The Netherlands, p. 79.
19. Ратников В. Ю. On the finds of Pliobatrachus (Anura, Palaeobatrachidae) in Eastern Europe. Paleintological Journal, 1997, vol. 31, no. 4, pp. 415–421.
20. Ratnikov V. Yu. Pliocene anurans of East-European platform. Russian journal of herpetology, 2001, vol.8, no 3, pp. 171–178.
21. Ratnikov V. Yu. Muchkapian (Early Neopleistocene) Amphibians and Reptiles of the East-European plain. Russian journal of herpetology, 2002, vol.9, no. 3, P. 229–236.
22. Ratnikov V. Yu. Novye nakhodki zemnovodnykh i и presmykayushchikhsya v opornykh muchkapskikh mestonakhozhdeniyakh basseina Verkhnego Dona [New finds of amphibians and reptiles in the reference Muchkap localities of the Upper Don basin]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya = Proceedings of Voronezh State University. Series: Geology, 2002, no. 1, pp. 73–79. (In Russ.)
23. Ratnikov V. Yu. Novye nakhodki zemnovodnykh i presmykayushchikhsya iz nizhneneopleistotsenovogo mestonakhozhdeniya Il'inka [New Finds of Amphibians and Reptiles from the Lower Neopleistocene Ilyinka Locality]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya = Proceedings of Voronezh State University. Series: Geology, 2003, no. 1, pp. 36–39. (In Russ.)
24. Ratnikov V. Yu. Osteology of Russian toads and frogs for palaeontological researches. Acta zool. cracov., 2001, vol. 44, no. 1, pp. 1–23.
25. Ratnikov V. Yu. Osteologicheskie kharakteristiki nadvidovykh taksonov zhab i lyagushek Vostochnoi Evropy [Osteological characteristics of supraspecific taxa of toads and frogs of Eastern Europe] Voprosy gerpetologii: materialy pyatogo s'ezda Gerpetologicheskogo obshchestva im. A.M. Nikol'ckogo [Questions of herpetology: materials of the Fifth Congress of the Herpetological Society named after A.M. Nikolsky], Minsk, September 25-28, 2012, pp. 269–273. (In Russ.)
26. Čerňansky ́ A., Syromyatnikova E. V. The first Miocene fossils of Lacerta cf. trilineata (Squamata, Lacertidae) with a comparative study of the main cranial osteological differences in green lizards and their relatives. PLoS ONE, 2019, vol.14, no. 8: e0216191, pp. 1–55.
27. Vorob'eva E. I., Chugunova T. Yu. Zubnaya sistema yashcherits: taksonomicheskoe i ekologicheskoe raznoobrazie. [Dental system of lizards: taxonomic and ecological diversity]. Moscow, Nauka, 1995, 151 p. (In Russ.)
28. Goncharov A. G. O rasprostranenii yashcherits (Sauria) v Tsentral'nom Chernozem'e. [About the distribution of lizards (Sauria) in the Central Black Earth Region]. Sovremennaya gerpetologiya: problemy i puti ikh resheniya. Stat'i po materialam dokladov Pervoi mezhdunarodnoi molodezhnoi konferentsii gerpetologov Rossii i sopredel'nykh stran. [Modern herpetology: problems and solutions. Articles based on the materials of the reports of the First International Youth Conference of Herpetologists in Russia and neighboring countries]. Saint-Peterburg, Russia, November 25–27, 2013, pp. 72–76. (In Russ.)
29. Ratnikov V. Yu. Gerpetofauna iz sredneneopleistotsenovogo mestonakhozhdeniya Ruchei Kolabinsky (Kursraya oblast'). [Herpetofauna from the Late Middle Pleistoсene locality of Kolabinsky brook (Kursk region). Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya = Proceedings of Voronezh State University. Series: Geology, 2019, no. 2, pp. 36–42. (In Russ.)
30. Szyndlar Z. A review of Neogene and Quaternary snakes of Central and Eastern Europe. Part II: Natricinae, Elapidae, Viperidae. Estudios geol., 1991, vol. 47, pp. 237–266.
31. Auffenberg W. The fossil Snakes of Florida. Tulane Studies Zool., 1963, vol. 10, no 3, pp. 131–216.
32. Ratnikov V. Yu. Identification of some Eurasian species of Elaphe (Colubridae, Serpentes) on the basis of vertebrae. Russian Journal of Herpetology, 2004, vol. 11, no. 2, pp. 91–98.
33. Ratnikov V., Mebert K. Fossil remains of Natrix tessellata from the Late Cenozoic deposits of the East European Plain. Mertensiella, 2011, no. 18, pp. 337–342.
34. Ananjeva N. B., Orlov N. L., Khalikov R. G., Darevsky I. S., Ryabov S. A., Barabanov A. V. The Reptiles of Northern Eurasia (Taxonomic Diversity, Distribution, Conservation Status). Saint-Petersburg, Zoological Institute RAS, 2006, 245 p.
35. Ivanov M. The first European pit viper from the Miocene of Ukraine. Acta Palaeontologica Polonica, 1999, vol. 44, no. 3, pp. 327–334.
36. Ratnikov V. Yu. Gerpetofauna nizhnepleistotsenovogo mestonakhozhdeniya Vol'naya Vershina v basseine Verkhnego Dona [Herpetofauna of the Lower Pleistocene locality Volnaya Vershina in the Upper Don basin]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya = Proceedings of Voronezh State University. Series: Geology, 1996, no. 1, pp. 90–93. (In Russ.)
37. Ratnikov V. Yu. Eopleistocene and Pleistocene Anurans of the East-European Platform. Paleont. Zhurn., 1992, no. 1, pp. 89– 100. (In Russ.)
38. Ratnikov V. Yu. Novye dannye o gerpetofaune mestonakhozhdeniya Kuznetsovka v Tambovskoi oblasti. [New data on the herpetofauna of the Kuznetsovka locality in the Tambov region]. Izv. vuzov. Geologiya i razvedka = Proceedings of universities. Geology and exploration, 1997, no. 1, pp. 26–32. (In Russ.)
39. Ratnikov V. Yu. Fossil remains of modern amphibian and reptile species as the material for studying of their areas history. Trudy Nauchno-issledovatel'skogo Instituta Geologii [The work of the Research Institute of Geology], Voronezh, VSU Publ., vol. 59, 2009, 91 p. (In Russ.).
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Опубликован
2020-12-01
Как цитировать
Ратников, В. Ю. (2020). Плио-плейстоценовые герпетофауны окрестностей села Коротояк (Воронежская область). Вестник ВГУ. Серия: Геология, (4), 12-25. https://doi.org/10.17308/geology.2020.4/3123
Раздел
Палеонтология, литология, стратиграфия
