Новые данные об эоплейстоценовой герпетофауне из местонахождения Морозовка-1 (Одесская область)
Ключевые слова:
эоплейстоцен, герпетофауна, амфибии, рептилии, палеогеография
Аннотация
Введение: герпетофауна из эоплейстоценового местонахождения Морозовка-1 уже изучалась ранее. Из-за плохой сохранности ископаемых остатков и недостатка сравнительного материала до вида удалось определить лишь три змеиных позвонка из семи. Недавно при разборе коллекций были обнаружены еще два змеиных позвонка. Малочисленность изученных материалов по эоплейстоценовым герпетофаунам Восочной Европы делает необходимым уточнить прежние определения и доопределить неизученные ранее позвонки змей из этого местонахождения.
Методика исследований: для видовой идентификации ископаемые позвонки сравнивались с позвонками современных видов.
Систематическая часть: приводится описание новых и переописание ранее изучавшихся позвонков змей. Они определены как Coronella austriaca, Elaphe sauromates, Elaphe dione, Colubrinae indet., Colubridae indet., Vipera berus, Vipera ursinii s.l.
Обсуждение и выводы: с учетом ранее изученных материалов остатки амфибий и рептилий из местонахождения Морозовка-1 (Черевичное-1) представлены следующими формами: слой 4: Pelobates fuscus aut vespertinus – 1, Pelobatidae indet. – 1, Anura indet. – 1; Testudines indet. – 1, Lacerta agilis L. – 1, Elaphe dione – 1, Colubrinae indet. – 1, Natrix natrix (L.) – 2, Natrix tessellata (Laur.) – 1, Colubridae indet. – 1, Vipera berus – 1, Vipera ursinii s.l. – 1; слой 5: Coronella austriaca – 1, Elaphe sauromates – 1. По количественному соотношению остатков видов, предпочитающих различные биотопы, можно предположить лесостепную обстановку с участием широколиственных и смешанных лесов, чередующихся с открытыми пространствами и редколесьями. Климат, скорее всего, был близок к современному в районе местонахождения.
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Литература
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12. Ratnikov V., Mebert K. Fossil remains of Natrix tessellata from the Late Cenozoic deposits of the East European Plain. Mertensiella, 2011, no. 18, pp. 337–342
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19. Ratnikov V. Yu. Pozdnekajnozojskie zemnovodnye i cheshujchatye presmykajushhiesja Vostochno-Evropejskoj ravniny [Late Cenozoic amphibians and reptiles of the East-European plain]. Trudy nauchnoissledovatel'skogo instituta geologii [The work of the Research Institute of Geology], Voronezh: VSU publ, vol. 10, 138 p. (In Russ.)
20. Ratnikov V. Yu. Obzor pozdnekajnozojskikh ostatkov yashcherits Vostochno-Evropejskoj ravniny [A review of the late Cenozoic lizard remains of the East European Plain]. Samarskaja Luka: problemy regional’noj I global’noj ekologii − Samara Luka: problems of regional and global ecology, 2009, vol. 18, no. 1, pp. 111-115 (In Russ.)
21. Ratnikov V. Yu. Pozdneneopleistotsenovaja gerpetofauna iz mestonakhozhdenija Staryj Oskol (Belgorodskaja oblast’) [Late Neopleistocene herpetofauna from the Stary Oskol locality (Belgorod region)]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya − Proceedings of Voronezh State University. Series: Geology, 2019, no. 4, pp. 31–36 (In Russ.)
22. Ratnikov V. Yu. Plio-Pleistotsenovye gerpetofauny okrestnostej sela Korotojak (Voronezhskaja oblast’) [Plio-Pleistocene herpetofaunas of the vicinity of the village of Korotoyak (Voronezh region)]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya − Proceedings of Voronezh State University. Series: Geology, 2020, no. 4, pp. 12–26 (In Russ.)
23. Ratnikov V. Yu., Motuzko A. N., Akimova E. V. Herpetofauna i melkije mlekopitajushchie pozdnepaleoliticheskoj stojan¬ki Listvenka [Herpetofauna and small mammals of the Late Paleolithic camp Listvenka]. Problemy arkheologii, etnografii, antropologii Sibiri i sopredel’nykh territorij − Problems of archeology, ethnography, anthropology of Siberia and adjacent territories, Novosibirsk, 2009, pp. 186–189 (In Russ.)
24. Ratnikov V. Yu. Muchkapian (Early Neopleistocene) Amphibians and Reptiles of the East-European plain. Russian journal of herpetology, 2002, vol. 9, no. 3, pp. 229–236
25. Zerova G. A., Chkhikvadze V. M. Obzor kajnozojskikh yashcherits i zmej SSSR [Review of Cenozoic lizards and snakes of the USSR]. Izvestija Akademii nauk GSSR − Serija biologicheskaja, 1984, vol. 10, no. 5, pp. 319–326 (In Russ.)
26. Gutieva N. V., Chkhikvadze V. M. Gigantskaja ljagushka iz pleistotsena peshchery Okladnikova (Gornyj Altaj) [Giant frog from the Pleistocene of Okladnikov Cave (Altai Mountains)]. Khronostratigrafija paleolita Severnoj, Tsentral’noj i Vostochnoj Azii i Ameriki: doklady mezhdunarodnogo simposiuma [Chronostratigraphy of the Paleolithic of Northern, Central and Eastern Asia and America: proceedings of the International Symposium]. Novosibirsk publ., 1990. pp. 132–134 (In Russ.)
27. Ratnikov V. Yu., Maslova I. V., Omelko V. E., Tiunov M. P. (2024) Cheshujchatyje presmykajushchiesja pozdnego golotsena iz otlozhenij peshchery Medvezhij Klyk na khrebte Lozovyj (Juzhnyj Sikhote-Alin’) [Late Holocene squamated reptiles from the Medvezhiy Klyk Cave on the Lozovy Ridge (Southern Sikhote-Alin, Primorsky Kray)]. Amurskij zoologicheskij zhurnal − Amurian Zoological Journal, vol. 16, no. 3, pp. 763–781 (In Russ.)
28. Jakovleva T. I. Golotsenovyje nakhodki zemnovodnykh i presmykajushchikhsja, vkljuchennykh v Krasnuju knigu Respubliki Bashkortostan [Holocene finds of amphibians and reptiles included in the Red Book of the Republic of Bashkortostan]. Universitetskaja nauka – Respublike Bashkortostan: Tom 1. Estestvennyje nauki: materialy konferentsii [University Science – to the Republic of Bashkortostan: Volume 1. Natural Sciences: proceedings of the scientific and practical conference]. Ufa publ., 2004, pp. 159–161 (In Russ.)
29. Chen Y., Li Y. X., Shi J. S., Zhang Y. X., Xie K. Pleistocene fossil snakes (Squamata, Reptilia) from Shanyangzhai Cave, Hebei, China. Historical Biology, 2021, vol. 33, no. 5, pp. 699–711
30. Ratnikov V. Yu. Likhvinian (Middle Neopleistocene) amphibians and reptiles of the East-European Plain. Russian Journal of Herpetology, 2005, vol. 12, no. 1, pp. 7–12
31. Szyndlar Z. A review of Neogene and Quaternary snakes of Central and Eastern Europe. Part I: Scolecophidia, Boidae, Colubrinae. Estudios Geol., 1991, vol. 47, pp. 103–126
32. Ratnikov V. Yu. Cheshujchatyje presmykajushchiesja iz verkhneneopleistotsenovykh otlozhenij peshchery Emine-Bair-Khosar v Krymu [Squamated reptiles from the Upper Neopleistocene deposits of the Emine-Bair-Khosar cave in Crimea]. Biospeleologija Kavkaza i drugikh rajonov Rossii: materialy konferentsii [Biospeleology of the Caucasus and other regions of Russia: materials of the All-Russian youth conference]. Moscow publ., 2015, pp. 64–65 (In Russ.)
33. Blain H. A. Amphibians and squamate reptiles from Azokh 1. Azokh cave and the transcaucasian corridor, 2016, pp. 191–210
34. Redkozubov O. I. Neogenovyje reptilii Moldovy: avtoref. diss. … kand. biol. nauk. [Neogene reptiles of Moldova: PhD thesises], Moscow publ., 1991, 20 p. (In Russ.)
35. Ratnikov V. Yu. Gerpetofauna iz sredneneopleistotsenovogo mestonakhozhdenija Ruchej Koljabinskij (Kurskaja oblast’) [Herpetofauna from the Middle Neopleistocene locality of Ruchey Kolyabinsky (Kursk region)]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya − Proceedings of Voronezh State University. Series: Geology, 2019, no. 2, pp. 36–42 (In Russ.)
36. Ratnikov V. Yu., Ivanov D. L. Golotsenovaja gerpetofauna iz mestonakhozhdenija Zarech’e na Vikhre (Belarus’) [Holocene herpetofauna from the Zarechye na Vikhra locality (Belarus)]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya − Proceedings of Voronezh State University. Series: Geology, 2024, no. 2, pp. 19–28 (In Russ.)
37. Ratnikov V. Yu., Ivanov D. L. Golotsenovaja gerpetofauna iz mestonakhozhdenija “Pionerskij” (Belarus’) [Holocene herpetofauna from the “Pionerskij” locality (Belarus)]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya − Proceedings of Voronezh State University. Series: Geology, 2024, no. 1, pp. 66–73 (In Russ.)
38. Ratnikov V. Yu., Blain H-A. Holocene amphibians and reptiles from Voroncha (Belarus): Comparative osteology, paleopathology and paleobiogeography. Historical Biology, 2020, vol. 32, no. 4, pp. 508–527.
39. Ratnikov V. Yu. Dynamics of East European modern amphibian and reptile species distribution areas and their potential use in Quaternary stratigraphy. Comptes Rendus Palevol, 2016, vol. 15, no. 6, pp. 721–730.
2. Konstantinova N. A. Geologicheskie uslovija mestonakhozhdenija melkkh mlekopitajuschikh v eopleistotsene juzhnoj Moldavii I yugo-zapadnoj Ukrainy [Geological conditions of a locality of small Eopleistocene mammals in Southen Moldavia and South-Western Ukraine]. Stratigraficheskoe znachenie antropogenovoj fauny melkih mlekopitajushhih − Stratigraphic importance jf small mammalian Anthropogen fauna. Moscow, 1965, pp. 60–97 (In Russ.)
3. Ananjeva N. B., Orlov N. L., Khalikov R. G. The reptiles of Northern Eurasia: Taxonomic diversity, distribution, conservation status. Sofia, Pensoft publ., 2006. 247 p.
4. Dunaev E. A., Orlova V. F. Zemnovodnye I presmykayuschiesya Rossii: Atlas-opredelitel’ [Amphibians and reptiles of Russia: Atlas-identifier]. 2nd edition, revised and enlarged. Moscow, Phiton publ., 2017. 328 p. (In Russ.)
5. Holman J. A. Pleistocene amphibians and reptiles in Britain and Europe. New York and Oxford: Oxford University Press, 1998, 254 p.
6. Gilmore C.W. Fossil snakes of North America. Geological society of America, special papers, 1938, no. 9, 96 p.
7. Auffenberg W. The fossil Snakes of Florida. Tulane studies in zoology, 1963, vol. 10, no. 3, pp. 131–216
8. Szyndlar Z. Fossil snakes from Poland. Acta zool. Cracov, 1984, vol. 28, no. 1, pp. 1–156
9. Ratnikov V. Yu. Identification of some Eurasian species of Elaphe (Colubridae, Serpentes) on the basis of vertebrae. Russian Journal of Herpetology, 2004, vol. 11, no. 2, pp. 91–98
10. Ratnikov V. Yu. Comparative Osteology of Two Far Eastern Species of Ratsnakes (Serpentes: Colubridae), Elaphe dione (Pallas, 1773) and E. schrenckii (Strauch, 1873), for the Purpose of Palaeontological Studies. Asian Herpetol. Res., 2022, vol. 13, no. 1, pp. 1–22
11. Ratnikov V. Yu. Vertebrae of modern viperids of the Far East: Comparative characterisation for the identification of their fossil remains. Asian Herpetol Res., 2024, vol. 15, no. 3, pp. 152–172
12. Ratnikov V., Mebert K. Fossil remains of Natrix tessellata from the Late Cenozoic deposits of the East European Plain. Mertensiella, 2011, no. 18, pp. 337–342
13. Uetz P., Freed P., Aguilar R. The Reptile Database, 2025. Available at: http://www.reptile-database.org, (accessed 19.08.2025)
14. Szyndlar Z., Rage J-C. Oldest Fossil Vipers (Serpentes: Viperidae) from the Old World. Kaupia, 1999, vol. 8, pp. 9–20
15. Ivanov M., Čerňanský A. Vipera berus (Linnaeus, 1758) remains from the late Pleistocene of Slovakia. Amphibia-Reptilia, 2017, vol. 38, no. 2, pp. 133–144
16. Frost D. R. Amphibian Species of the World: an Online Reference. Version 6.2. 2025. Available at: https://amphibiansoftheworld.amnh.org/index.php. American Museum of Natural History, New York, USA. doi.org/10.5531/db.vz.0001 (accessed 19.08.2025)
17. Ratnikov V. Yu. Iskopaemye ostatki sovremennykh vidov zemnovodnykh i cheshujchatykh presmykajuschikhsja kak material dlja izuchenija istorii ikh arealov [Fossil remains of modern species of amphibians and squamated reptiles as material for studying the history of their areas]. Trudy Nauchno-issledovatel'skogo Instituta Geologii [The work of the Research Institute of Geology], Voronezh, VSU publ., vol. 59, 2009, 91 p. (In Russ.)
18. Ratnikov V. Yu. Novye nakhodki zemnovodnykh i presmykayuschikhsja v opornykh muchkapskikh mestonakhozhdenijakh basseina Verkhnego Dona [New finds of amphibians and reptiles in the key Muchkap localities of the Upper Don basin]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya − Proceedings of Voronezh State University. Series: Geology, 2002, no. 1, pp. 73–79 (In Russ.)
19. Ratnikov V. Yu. Pozdnekajnozojskie zemnovodnye i cheshujchatye presmykajushhiesja Vostochno-Evropejskoj ravniny [Late Cenozoic amphibians and reptiles of the East-European plain]. Trudy nauchnoissledovatel'skogo instituta geologii [The work of the Research Institute of Geology], Voronezh: VSU publ, vol. 10, 138 p. (In Russ.)
20. Ratnikov V. Yu. Obzor pozdnekajnozojskikh ostatkov yashcherits Vostochno-Evropejskoj ravniny [A review of the late Cenozoic lizard remains of the East European Plain]. Samarskaja Luka: problemy regional’noj I global’noj ekologii − Samara Luka: problems of regional and global ecology, 2009, vol. 18, no. 1, pp. 111-115 (In Russ.)
21. Ratnikov V. Yu. Pozdneneopleistotsenovaja gerpetofauna iz mestonakhozhdenija Staryj Oskol (Belgorodskaja oblast’) [Late Neopleistocene herpetofauna from the Stary Oskol locality (Belgorod region)]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya − Proceedings of Voronezh State University. Series: Geology, 2019, no. 4, pp. 31–36 (In Russ.)
22. Ratnikov V. Yu. Plio-Pleistotsenovye gerpetofauny okrestnostej sela Korotojak (Voronezhskaja oblast’) [Plio-Pleistocene herpetofaunas of the vicinity of the village of Korotoyak (Voronezh region)]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya − Proceedings of Voronezh State University. Series: Geology, 2020, no. 4, pp. 12–26 (In Russ.)
23. Ratnikov V. Yu., Motuzko A. N., Akimova E. V. Herpetofauna i melkije mlekopitajushchie pozdnepaleoliticheskoj stojan¬ki Listvenka [Herpetofauna and small mammals of the Late Paleolithic camp Listvenka]. Problemy arkheologii, etnografii, antropologii Sibiri i sopredel’nykh territorij − Problems of archeology, ethnography, anthropology of Siberia and adjacent territories, Novosibirsk, 2009, pp. 186–189 (In Russ.)
24. Ratnikov V. Yu. Muchkapian (Early Neopleistocene) Amphibians and Reptiles of the East-European plain. Russian journal of herpetology, 2002, vol. 9, no. 3, pp. 229–236
25. Zerova G. A., Chkhikvadze V. M. Obzor kajnozojskikh yashcherits i zmej SSSR [Review of Cenozoic lizards and snakes of the USSR]. Izvestija Akademii nauk GSSR − Serija biologicheskaja, 1984, vol. 10, no. 5, pp. 319–326 (In Russ.)
26. Gutieva N. V., Chkhikvadze V. M. Gigantskaja ljagushka iz pleistotsena peshchery Okladnikova (Gornyj Altaj) [Giant frog from the Pleistocene of Okladnikov Cave (Altai Mountains)]. Khronostratigrafija paleolita Severnoj, Tsentral’noj i Vostochnoj Azii i Ameriki: doklady mezhdunarodnogo simposiuma [Chronostratigraphy of the Paleolithic of Northern, Central and Eastern Asia and America: proceedings of the International Symposium]. Novosibirsk publ., 1990. pp. 132–134 (In Russ.)
27. Ratnikov V. Yu., Maslova I. V., Omelko V. E., Tiunov M. P. (2024) Cheshujchatyje presmykajushchiesja pozdnego golotsena iz otlozhenij peshchery Medvezhij Klyk na khrebte Lozovyj (Juzhnyj Sikhote-Alin’) [Late Holocene squamated reptiles from the Medvezhiy Klyk Cave on the Lozovy Ridge (Southern Sikhote-Alin, Primorsky Kray)]. Amurskij zoologicheskij zhurnal − Amurian Zoological Journal, vol. 16, no. 3, pp. 763–781 (In Russ.)
28. Jakovleva T. I. Golotsenovyje nakhodki zemnovodnykh i presmykajushchikhsja, vkljuchennykh v Krasnuju knigu Respubliki Bashkortostan [Holocene finds of amphibians and reptiles included in the Red Book of the Republic of Bashkortostan]. Universitetskaja nauka – Respublike Bashkortostan: Tom 1. Estestvennyje nauki: materialy konferentsii [University Science – to the Republic of Bashkortostan: Volume 1. Natural Sciences: proceedings of the scientific and practical conference]. Ufa publ., 2004, pp. 159–161 (In Russ.)
29. Chen Y., Li Y. X., Shi J. S., Zhang Y. X., Xie K. Pleistocene fossil snakes (Squamata, Reptilia) from Shanyangzhai Cave, Hebei, China. Historical Biology, 2021, vol. 33, no. 5, pp. 699–711
30. Ratnikov V. Yu. Likhvinian (Middle Neopleistocene) amphibians and reptiles of the East-European Plain. Russian Journal of Herpetology, 2005, vol. 12, no. 1, pp. 7–12
31. Szyndlar Z. A review of Neogene and Quaternary snakes of Central and Eastern Europe. Part I: Scolecophidia, Boidae, Colubrinae. Estudios Geol., 1991, vol. 47, pp. 103–126
32. Ratnikov V. Yu. Cheshujchatyje presmykajushchiesja iz verkhneneopleistotsenovykh otlozhenij peshchery Emine-Bair-Khosar v Krymu [Squamated reptiles from the Upper Neopleistocene deposits of the Emine-Bair-Khosar cave in Crimea]. Biospeleologija Kavkaza i drugikh rajonov Rossii: materialy konferentsii [Biospeleology of the Caucasus and other regions of Russia: materials of the All-Russian youth conference]. Moscow publ., 2015, pp. 64–65 (In Russ.)
33. Blain H. A. Amphibians and squamate reptiles from Azokh 1. Azokh cave and the transcaucasian corridor, 2016, pp. 191–210
34. Redkozubov O. I. Neogenovyje reptilii Moldovy: avtoref. diss. … kand. biol. nauk. [Neogene reptiles of Moldova: PhD thesises], Moscow publ., 1991, 20 p. (In Russ.)
35. Ratnikov V. Yu. Gerpetofauna iz sredneneopleistotsenovogo mestonakhozhdenija Ruchej Koljabinskij (Kurskaja oblast’) [Herpetofauna from the Middle Neopleistocene locality of Ruchey Kolyabinsky (Kursk region)]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya − Proceedings of Voronezh State University. Series: Geology, 2019, no. 2, pp. 36–42 (In Russ.)
36. Ratnikov V. Yu., Ivanov D. L. Golotsenovaja gerpetofauna iz mestonakhozhdenija Zarech’e na Vikhre (Belarus’) [Holocene herpetofauna from the Zarechye na Vikhra locality (Belarus)]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya − Proceedings of Voronezh State University. Series: Geology, 2024, no. 2, pp. 19–28 (In Russ.)
37. Ratnikov V. Yu., Ivanov D. L. Golotsenovaja gerpetofauna iz mestonakhozhdenija “Pionerskij” (Belarus’) [Holocene herpetofauna from the “Pionerskij” locality (Belarus)]. Vestnik Voronezhskogo gosudarstvennogo universiteta. Seriya: Geologiya − Proceedings of Voronezh State University. Series: Geology, 2024, no. 1, pp. 66–73 (In Russ.)
38. Ratnikov V. Yu., Blain H-A. Holocene amphibians and reptiles from Voroncha (Belarus): Comparative osteology, paleopathology and paleobiogeography. Historical Biology, 2020, vol. 32, no. 4, pp. 508–527.
39. Ratnikov V. Yu. Dynamics of East European modern amphibian and reptile species distribution areas and their potential use in Quaternary stratigraphy. Comptes Rendus Palevol, 2016, vol. 15, no. 6, pp. 721–730.
Опубликован
2025-12-31
Как цитировать
Ратников, В. Ю. (2025). Новые данные об эоплейстоценовой герпетофауне из местонахождения Морозовка-1 (Одесская область). Вестник ВГУ. Серия: Геология, (4), 34-44. https://doi.org/10.17308/geology/1609-0691/2025/4/34-44
Раздел
Палеонтология, литология, стратиграфия
